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by Jason A. Dunlop*1 Introduction: Arachnida is one of the major arthropod groups. It includes spiders (Araneae), scorpions (Scorpiones), mites (Acari) and harvestmen (Opiliones), as well as a number of rarer and less familiar groups (Fig 1). The name Arachnida was introduced by the French zoologist Jean-Baptise Lamarck and is derived from Greek mythology: in one story, the maiden Arachne challenged the goddess Athene to a weaving contest, and was subsequently transformed into a spider — condemned to weave for evermore. There are about 100,000 living species of arachnids, with mites and spiders representing the most diverse and species-rich groups. Fossil arachnids are considerably rarer, with more than 1,700 described species (well over half of which are spiders) and a record that exten...

by Jason A. Dunlop*1 Introduction: Chasmataspidida (Fig 1) are rare, extinct arthropods known only from the early to mid Palaeozoic Era. They are probably closely related to either xiphosurans (horseshoe crabs; Fig 2) or eurypterids (sea scorpions; Fig 1); some chasmataspid fossils were originally misinterpreted as members of one of these two groups. They were first discovered in the 1950s, and were only recently recognized as a group distinct from horseshoe crabs. Chasmataspids are currently the oldest known examples of the Euchelicerata lineage —all Chelicerata except Pycnogonida (sea spiders) — and palaeontologists date the origins of these euchelicerates back to at least the late Cambrian Period. Morphology: Most chasmataspids are a few centimetres long. The body is divided int...

by Jason A. Dunlop*1 Introduction: Pycnogonida, or sea spiders, are not true spiders at all. They are in fact a group of — probably rather primitive — marine arthropods, characterized by a small, slender body and in many cases by correspondingly long legs (Fig. 1). So unusual is their morphology that many of their internal-organ systems have been displaced into the legs. Because of their strange appearance, older studies occasionally referred to them as ‘nobody crabs’ (literally crabs without a body) — although it is important to stress that they are not crustaceans, any more than they are spiders. Pycnogonids are thought either to have evolved right at the very base of the arthropod tree — and thus not to be closely related to any particular group of arthropods — or to be related to ara...

by Jason A. Dunlop*1 Introduction: Chelicerata is one of the main divisions of the arthropods, and essentially consists of arachnids and their closest relatives. The name was coined in 1901 by the Berlin-based zoologist Richard Heymons (Fig. 1). It means the ‘claw-bearers’, in reference to the claw- or fang-shaped mouthparts that characterize the group. In addition to the arachnids, Chelicerata also includes the horseshoe crabs (Xiphosura), the extinct sea scorpions (Eurypterida) and little-known chasmatapaids (Chasmataspidida), and the sea spiders (Pycnogonida). The inclusion of sea spiders within this group is controversial, as we shall see below, and arachnids, horseshoe crabs, eurypterids and chasmataspids are sometimes grouped together as the Euchelicerata. The name Merostoma...